Plot
A secret laboratory near the Arctic Circle, is stolen by a mysterious vehicle using a giant magnet. The Anti-Villain League (AVL) become suspicious about this case as the lab contains a mutating chemical compound, PX-41, which could transform living things into indestructible killing machines. The organization decides to recruit Gru, as he is a former villain, and could help the AVL prevent the mutating chemical from getting into the wrong hands.
Lucy Wilde, an AVL agent forcibly abducts Gru after stunning him unconscious with her lipstick taser. Two of Gru's minions who try to thwart her plans are captured also. They travel to AVL's underwater headquarters where Gru meets the league's director, Silas Ramsbottom. He asks Gru to assist them in their efforts to track down the perpetrator and the missing mutagen. Gru refuses, preferring to concentrate on his new venture of producing bottled jellies and the responsibilities of being a father to his three adopted girls, Margo, Agnes and Edith. Before he leaves, Lucy admits she was impressed by his work as a villain and gives her business card to Gru, urging him to contact her if he changes his mind. Upon return to his lab, Gru realises that his minion-run production line had churned out bottles of awful-tasting jellies. Also, his research scientist, Dr. Nefario confesses to Gru that he misses being "evil" and has been offered employment elsewhere. After giving Dr. Nefario a 21 fart-gun send-off with the minions, Gru contacts Lucy and takes on the task of recovering the stolen mutagen.
Traces of chemical scent given off by the mutagen narrow the search to the confines of The Paradise Shopping Mall. Gru works undercover as the owner of a cupcake shop in that mall with Lucy being assigned as his partner. He suspects one of the owners of a restaurant in the mall, named Eduardo, is an old super-villain called El Macho, who supposedly died after body-strapping TNT and riding a shark into an active volcano that resulted in his fiery demise. Lucy and Gru then decide to break into his restaurant but are spotted by Eduardo's pet chicken and narrowly escape from being caught by Eduardo when he returns back to his shop late in the night. However, they find only a jar of secret-recipe salsa in his safe and nothing relevant to link Eduardo to the crime. While the team investigates wig merchant Floyd Eagle-san, Margo begins to develop a crush on Eduardo's son, Antonio, to Gru's frustration. The whole family is invited to Eduardo's Cinco de Mayo party at his mansion where Gru's protectiveness of his daughter leads him to make baseless wild accusations to the AVL's Silas about Eduardo as El Macho and his son, which are scoffed at.
Later that day, a determined matchmaking neighbour, Jillian, sets a reluctant Gru up on a date with her rude and superficial friend Shannon. During the date, Shannon discovers Gru's wig and threatens to humiliate him, but Lucy saves him from embarrassment by shooting her with a tranquilizing dart. They take Shannon home, and Gru realizes he is in love with Lucy. When he arrives at the mall the next day, the Anti-Villain League arrest Floyd Eagle-san, who protests vehemently that he was framed, after the AVL team uncovers the empty mutagen jar in his shop. Silas tells Gru that the case is now closed and that Lucy will be assigned to Australia for work. Gru, heartbroken from the situation, tells Agnes that she was right that he loves her. However, Gru is unable to muster the courage to ask Lucy out before she leaves.
The family then goes to Eduardo's party. Gru follows Eduardo and discovers that he was correct about Eduardo being the presumed-dead super-villain El Macho. Gru also discovers that Dr. Nefario has been working for Eduardo and that the duo has been capturing and mutating some of Gru's minions into the purple-furred monsters using the stolen mutagen serum. Eduardo offers Gru a chance to conquer the world, but Gru makes unconvincing excuses and hastily leaves with the girls. Margo has also broken up with Antonio as he became interested in another girl. For breaking her heart, Gru encases Antonio in a block of ice with his freeze-ray.
Lucy, while in the plane bound for Australia, realizes that she wants to be with Gru, and jumps out to hang-glide to the party. Eduardo discovers her true identity and captures her after his chicken retrieves her AVL ID badge from her purse. Dr. Nefario calls Gru to tell him that Lucy has been captured. Gru disguises two of his minions in purple paint to escort his "captured" self into Eduardo's compound in a rescue attempt. Gru manages to restore the minions who were under El Macho's control using the awful-tasting jam infused with the antidote which Dr. Nefario made and defeats El Macho. After El Macho collapses to the ground, Gru tries to untie Lucy who is strapped onto a TNT-loaded shark rocket. While attempting to do so, Eduardo's pet chicken triggers the launch remote, sending the pair flying towards the lava-spewing volcano where El Macho faked his death. While on the rocket, Lucy accepts Gru's date invitation before the pair dives into the safety of the ocean, just seconds before the rocket explodes. One hundred and forty-seven dates later, the two are married and the girls finally have a mother. The minions close with a rendition of "Y.M.C.A" and "I Swear" (mistakenly sung as "Underwear") while the family dances. However, soon after the final picture is taken, a mutated purple furry minion appears, surprising the family.
During the credits, three Minions are seen "auditioning" for the upcoming Minions movie. However, a black wall showing the credits keeps falling and blocking their way.
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